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dc.contributor.advisor Watson, Maxine en Bogonovich, Marc David en 2013-05-16T01:26:00Z en 2013-05-16T01:26:00Z en 2013-05-15 en 2012 en
dc.identifier.uri en
dc.description Thesis (Ph.D.) - Indiana University, Biology, 2012 en
dc.description.abstract In this dissertation I explore broad-scale regularities in the distribution of North American ferns and lycophytes. I test models for the evolution of these regularities. To this end, I construct a comprehensive database of all North American fern and lycophyte species. In the final empirical chapter I extend my analyses to New World mammals and global amphibians. First, I describe patterns of North American fern and lycophyte species and family richness and describe the relationships of these patterns with climate. Next, I explore the geography of fern polyploid species and present biogeographical evidence that most polyploid speciation events are evolutionary dead-ends. These results contrast with notions that polyploid speciation leads to evolutionary novelty and adaptive geographical expansion. Next, I test the "Out of The Tropics" model (OTT) for the evolution of latitudinal gradients in diversity. The OTT model proposes that higher taxa tend to originate in the tropics, and subsequently expand into temperate regions while maintaining a tropical presence. The OTT model predicts that older and diverse higher taxa will have expanded from their region of tropical origin, while younger and less diverse higher taxa will still be located in the tropics. I find partial support for the OTT model. Endemic fern families are found only in the tropics suggesting fern families more frequently originate in the tropics than the extra-tropics. Fern families with many species extend further into temperate regions than do less diverse families. However, family age is not correlated with the degree of extension from the tropics. In the final empirical chapter, I describe a global biogeographical rule observed in ferns, mammals and amphibians. The new rule states that diversity of a taxon at the taxon's richness peak predicts the amount of geographical expansion of that taxon from its center, and the number of species completely outside the taxon's center. I call these patterns diversity-expansion relationships, and I explore these patterns using multiple measures of diversity and geographical breadth (expansion). I propose alternative explanations for the existence of diversity-expansion patterns and discuss their macroevolutionary implications. en
dc.language.iso en en
dc.publisher [Bloomington, Ind.] : Indiana University en
dc.rights Attribution 3.0 Unported (CC BY 3.0) en
dc.rights.uri en
dc.subject biogeography en
dc.subject diversity-expansion en
dc.subject fern en
dc.subject macroevolution en
dc.subject out of the tropics en
dc.subject polyploid en
dc.subject.classification Biology en
dc.subject.classification Evolution & development en
dc.subject.classification Ecology en
dc.title Broad-scale geographical evolution of ferns en
dc.type Doctoral Dissertation en

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